Study unit
3513123 Nutrient dynamics in forest ecosystems, 4 Cp  
Code 3513123  Validity 30.01.2013 -
Name Nutrient dynamics in forest ecosystems  Abbreviation MeRaDy 
Credits 4 Cp Date of expiry  
TypeAdvanced Studies Subject0951 Forestry 
ClassStudy Unit  Hours  
Study right   Grading0-5 
Recommended scheduling 
Organisation Forest Sciences (J) 

Study unit's general literature:
Author Title Year Obligatory library
     Binkley, D. & Fisher, R.F.  Ecology and Management of Forest Soils  2013  Obligatory library

Learning outcomes 

Advanced knowledge about nutrient cycling in forest ecosystems: inputs, outputs, and processes within the ecosystem.


Weathering and nitrogen fixation, nutrient retention in soil, reactions of different nutrients in soil, the role of soil organisms in the nutrient cycle, nutrient uptake and the function of mycorrhizas, the functions of nutrients in plants, litter formation and decomposition. Leaching and erosion. Impacts of environmental factors, fertilization, forest management, chemical precipitation and global change.

Modes of study 

Lectures (6 h), guided independent written work based on literature, oral presentation and attendance in 10 h of student presentations, literature, examination.

Study materials 

Binkley, D. & Fisher, R.F. Ecology and Management of Forest Soils. 2013. 304 p.

Evaluation criteria 

Attendance obligatory. Examination 0-5.


Senior Researcher in Forest Soil Science


Previous studies: Forest soils or corresponding studies.


The course is held every other year, next time in the spring term 2022.



Further information 

In English.


Letter (J, K) in front of the name of the course/exam indicates the campus on which teaching or exam takes place: J = Joensuu, K = Kuopio.

Present and future teaching
No teaching

Future exams
Functions Name Type Cp Teacher Timetable
Registration ended Nutrient dynamics in forest ecosystems  General examination   
03.06.22fri 12.00-16.00